Aradinae (Hemiptera, Heteroptera, Pentatomomorpha, Aradoidea, Aradidae – “flat bugs”) is mostly Holarctic (temperate and subtropics of Eurasia and North America) group. Here I have tried to summarize information about the taxonomy of the subfamily.
The most important catalogs are:
Berend Aukema (ed.). Catalogue of Palaearctic Heteroptera. 1995-2013 (the electronic database is supplemented to date). Available online:
https://catpalhet.linnaeus.naturalis.nl/linnaeus_ng/app/views/introduction/topic.php?id=9
Thomas J. Henry, Richard C. Froeschner (eds.). 1988 (eBook Published 17 July 2019). Catalog of the Heteroptera or True Bugs, of Canada and the Continental United States. DOI: https://doi.org/10.1201/9781351070447
María del Carmen Coscarón, Eugenia Fernanda Contreras. 2012. Catalog of Aradidae (Hemiptera: Heteroptera) for the Neotropical Region. DOI: https://doi.org/10.11646/zootaxa.3466.1.1
J. G. Pendergrast. 1968. The Aradidae of New Zealand (Hemiptera, Heteroptera) III – The Aradinae and Calisiinae. Transactions of the Royal Society of New Zealand: Zoology, Volume 10, Issue 10. Available online:
https://paperspast.natlib.govt.nz/periodicals/TRSZOO19680122.2.2
Aradinae – Atlas of Living Australia.
https://bie.ala.org.au/species/https://biodiversity.org.au/afd/taxa/ef3f3b71-284d-4b4b-9bcc-58646dc5f026#classification
Some identification guides (continental or country level, most of them somewhat outdated):
Heiss, E. & J. Péricart. 2007. Hemiptères Aradidae Piesmatidae et Dipsocoromorphes euro-méditerranéens. Faune de France n° 91. ISBN: 978-2-9030-5229-4 (Western Palearctic)
Ryuichi Matsuda. 1977. The Insects and Arachnids of Canada, part 3. The Aradidae of Canada. Available online: https://publications.gc.ca/site/eng/9.811337/publication.html (Canada
and adjacent states of the US)
J. R. de la Torre-Bueno. 1939. A Synopsis of the Hemiptera Heteroptera of America North of Mexico – Part I – Families Scutelleridae, Cydnidae, Pentatomidae, Aradidae, Dysodiiclae and Termitaphididae. Available online:
https://www.biodiversitylibrary.org/item/205520#page/276/mode/1up (the US and Canada)
Hsiao, T.Y., S.Z. Ren, L.Y. Zheng, H.L. Jing, H.G. Zou & S.L. Liu. 1981. A handbook for the determination of the Chinese Hemiptera-Heteroptera. Volume II: i-iv, 1-654 (China)
Y Nakatani, K Kohno, T Ishikawa, Mikio Takai, Tomohide Yasunaga, Ernst Heiss. 2012. Terrestrial Heteropterans: A Field Guide to Japanese Bugs, Volume 3. ISBN: 9784881371688 (Japan)
Wonwoong Kim, Minsuk Oh, WonGun Kim, Seunghwan Lee. 2023. New records of genus Aradus Fabricius (Hemiptera: Aradidae) from Korea, with first male description of Aradus gretae Kiritshenko. Journal of Asia-Pacific Biodiversity, Volume 16, Issue 2. Pages 157-162. ISSN 2287-884X, https://doi.org/10.1016/j.japb.2022.12.004. (https://www.sciencedirect.com/science/article/pii/S2287884X22001352) (Korea)
In the following preliminary list of species of Aradinae of the World fauna I may have missed some tropical species. The status of some taxa remains debatable. The list is available as a table (https://docs.google.com/spreadsheets/d/1QdLbUQZ9rm_jaDVs4YACV3_PD7gDrCujg5U8JEl2CUw/edit?usp=sharing). Unfortunately, only some Palaearctic species may be referred to “species groups” (according to Heiss & Péricart, 2007).
SUBFAMILY ARADINAE
Genus Aradiolus Kormilev 1967 (Neotropic)
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chemsaki Kormilev, 1980
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paradoxus Kormilev, 1967
Genus Aradus Fabricius, 1803 (World)
Subgenus Aradus s.str. (World)
Species known from Western Palaearctic
betulae group
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betulae (Linnaeus, 1758)
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brenskei Reuter, 1884
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caucasicus Kolenati, 1857
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elburzanus Heiss, 2004
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hieroglyphicus J. Sahlberg, 1878
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krueperi Reuter, 1884
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maghrebinus Heiss, 2003
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obscurus Vásárhelyi, 1988
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ribauti Wagner, 1956
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seidenstueckeri Heiss, 1989
pictus group
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annulicornis Fabricius, 1803
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horvathi Vásárhelyi, 1984
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obtectus Vásárhelyi, 1988
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pictus Baerensprung, 1859
corticalis group
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betulinus Fallén, 1807
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brevicollis Fallén, 1807
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corticalis (Linnaeus, 1758)
conspicuus group
- cilicicus Heiss, 1990
- conspicuus Herrich-Schaeffer, 1835
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inopinus Kiritshenko, 1955
erosus group
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erosus Fallén, 1807
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montandoni Reuter, 1885
lugubris group
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angularis J. Sahlberg, 1886
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aterrimus Fieber, 1864
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bimaculatus Reuter, 1872
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canariensis Kormilev, 1954
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laeviusculus Reuter, 1875
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lugubris Fallén, 1807
depressus group
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depressus (Fabricius, 1794) (including depressus depressus (Fabricius, 1794) and depressus leptocerus Horváth, 1882)
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versicolor Herrich-Schaeffer, 1835
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serbicus Horváth, 1888
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somcheticus Kiritshenko, 1913
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truncatus Fieber, 1860
mirus group
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mirus Bergroth, 1894
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reuterianus Puton, 1875
stenopterus group
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cinnamomeus Panzer, 1806
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graecus Heiss, 1997
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lauri Noualhier, 1893
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safavii Hoberlandt, 1974
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stenopterus Bergroth, 1887
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tauricus Jakovlev, 1907
species incertae sedis
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crenaticollis R.F. Sahlberg, 1848
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distinctus Fieber, 1860
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flavicornis Dalman, 1823
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kuthyi Horváth, 1899
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pallescens Herrich-Schaeffer, 1840 (including pallescens pallescens Herrich-Schaeffer, 1840 and pallescens frigidus Kiritshenko, 1913)
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rysakovi Kiritshenko, 1951
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signaticornis R.F. Sahlberg, 1848
Species known from Eastern Palaearctic only
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bergrothianus Kiritshenko, 1913
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candidatus Bergroth, 1889
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compar Kiritshenko, 1913
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consentaneus Horváth, 1905
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czerskii Kiritshenko, 1915
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dembickyi Heiss, 2007
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discompar Hsiao, 1964
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dissimilis A. Costa, 1847
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dissors Kiritshenko, 1913
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emeiensis Heiss, 2008
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esakii Kormilev & Heiss, 1976
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formosanus Heiss, 2009
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gaoligongensis Bai, Heiss & Cai, 2010
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gretae Kiritshenko, 1955
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herculeanus Kiritshenko, 1913
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holzschuhi Heiss, 2003
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honshuensis Heiss & Shono, 2005
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komarowii Jakovlev, 1885
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kyrgyzstanus Heiss, 2004
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miyamotoi Heiss & Shono, 2003
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nipponicus Kormilev, 1955
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omeiensis Hsiao, 1964
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orientalis Bergroth, 1885
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oroszi Vásárhelyi, 2016
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parayunnanus Heiss, 2010
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pictellus Kerzhner, 1973
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poppiusi Kiritshenko, 1913
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pulchellus J. Sahlberg, 1878
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quinlingshanensis Heiss, 2003
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semilacer Kiritshenko, 1913
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setiger Kiritshenko, 1913
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sinensis Kormilev, 1955
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smetanai Heiss, 2003
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spinicollis Jakovlev, 1880
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tonkinensis Kormilev & Heiss, 1976
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transiens Kiritshenko, 1913
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trapezonotus Heiss & Shono, 2005
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turnai Heiss, 2008
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unicolor Kiritshenko, 1913
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ussurensis Jakovlev, 1876
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vitiosus Jakovlev, 1889
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yunnanus Heiss, 2003
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zhengi Heiss, 2001
Species known from South Asia only (excl. candidatus (also Eastern Palaearctic))
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abnormis Bergroth, 1889
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burmensis Kormilev, 1976
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malaisei Kormilev, 1976
Species known from Nearctic (the US and Canada) (excl. Holarctic lugubris)
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abbas Bergroth, 1889
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acutus Say, 1832
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aequalis Say, 1832
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alaskanus Kormilev and Heiss, 1979
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ampliatus Uhler, 1876
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antennalis Parshley, 1921
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apicalis Van Duzee, 1920
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approximatus Parshley, 1921
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arizonicus Parshley, 1921
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barberi Kormilev, 1966
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basalis Parshley, 1921
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behrensi Bergroth, 1886
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blaisdelli Van Duzee, 1920
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borealis Heidemann, 1909
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breviatus Bergroth, 1887
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brevicornis Kormilev, 1980
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brunnicornis Blatchley, 1926
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carolinensis Kormilev, 1964
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cincticornis Bergroth, 1906
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coarctatus Heidemann, 1907
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coloradensis Kormilev, 1964
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compressus Heidemann, 1907
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concinnus Bergroth, 1892
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consors Parshley, 1921
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crenatus Say, 1832
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curticollis Bergroth, 1913
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debilis Uhler, 1876
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depictus Van Duzee, 1917
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duzeei Bergroth, 1892
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evermanni Van Duzee, 1920
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falleni Stål, 1860
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funestus Bergroth, 1913
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urnissi Usinger, 1936
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furvus Parshley, 1921
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fuscipennis Usinger, 1936
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fuscomaculatus Stål, 1859
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gracilicornis Stål, 1873
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gracilis Parshley, 1929
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hesperius Parshley, 1921
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implanus Parshley, 1921
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inornatus Uhler, 1876
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insignitus Parshley, 1921
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insolitus Van Duzee, 1916
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intectus Parshley, 1921
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intermedius Usinger, 1936
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kormilevi Heiss, 1980
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lawrencei Kormilev, 1966
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leachi Van Duzee, 1929
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linsleyi Usinger, 1936
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marginatus Uhler, 1893
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medioximus Parshley, 1921
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montanus Bergroth, 1913
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occidentalis Kormilev, 1980
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opertaneus Parshley, 1921
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orbiculus Van Duzee, 1920
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oregonicus Kormilev, 1978
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ornatus Say, 1832
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ovatus Kormilev, 1966
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oviventris Kormilev, 1966
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paganicus Parshley, 1929
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pannosus Van Duzee, 1920 (including pannosus pannosus Van Duzee, 1920 and pannosus incomptus Parshley, 1921)
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parshleyi Van Duzee, 1920
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parvicornis Parshley, 1921
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patibulus Van Duzee, 1927
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persimilis Van Duzee, 1916
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proboscideus Walker, 1873
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quadrilineatus Say, 1825
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robustus Uhler, 1871 (including robustus robustus Uhler, 1871 and robustus insignis Parshley, 1921)
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saileri Kormilev, 1966
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saskatchewanensis Matsuda, 1980
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serratus Usinger, 1936
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shermani Heidemann, 1907
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signaticornis Sahlberg, 1848
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similis Say, 1832
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snowi Van Duzee, 1920
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subruficeps Hussey, 1953
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taylori Van Duzee, 1920
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tuberculifer Kirby, 1837
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uniannulatus Parshley, 1921
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uniformis Heidemann, 1904
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vadosus Van Duzee, 1920
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vandykei Van Duzee, 1927
Species known from Neotropics only (excl. falleni and gracilicornis that are also occur in the US)
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angustellus (Blanchard, 1852)
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brasiliensis Usinger, 1940
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compressicornis Stål, 1873
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fronterana Drake, 1942
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granuliger Kormilev, 1980
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mexicanus Usinger, 1936
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pallidicornis Stål, 1873
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penningtoni Drake, 1942
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pericarti Heiss, 1994
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testaceus Kormilev, 1980
Species known from Australia and New Zealand
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albicornis (Walker, 1873)
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australis Erichson, 1842
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erraticus Kormilev, 1966
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fuscicornis Kormilev, 1966
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leucotelus (Walker, 1873)
Species known from Sub-Saharan Africa only
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gracilipes Linnavuori, 1978
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zavattarii Mancini,1939
Subgenus Lobataradus Heiss, 2008 (Estern Palaearctic, South Asia)
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bilobatus Heiss & Shono, 2005
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khuntanus Heiss, 2015
species inquirendae (type specimens are lost, maybe not Aradidae)
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griseus (Fabricius, 1794) (Acanthia grisea Fabricius, 1794)
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nigricornis (Fabricius, 1794) (Acanthia nigricornis Fabricius, 1794)
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tridentatus (Geoffroy, 1785) (Cimex tridentatus Geoffroy in Fourcroy, 1785)
Genus Miraradus Vásárhelyi, 1980 (Estern Palaearctic, South Asia)
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angulatus Heiss, 2009
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foliaceus (Kormilev, 1957)
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mirabilis (Bergroth, 1892)
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oervendetes Vásárhelyi, 1980
Genus Quilnus Stål, 1873 (Holarctic)
Species known from Western Palaearctic
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cedri (Puton, 1873)
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discedens (Horváth, 1911)
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marcosi Heiss & Baena, 2006
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mediterraneus Heiss, 1989
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parvicollis (Stål, 1873)
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subsimilis (Horváth, 1911)
Species known from Estern Palaearctic
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brevirostris (Horváth, 1901)
Species known from Nearctic
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heidemanni Bergroth, 1906
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niger Stål, 1873
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nigrinus Parshley, 1921 (including nigrinus nigrinus Parshley, 1921 and nigrinus canadensis Parshley, 1929)
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usingeri Kormilev, 1978
Publicado el
septiembre 26, 2022 06:56 MAÑANA
por
kgrebennikov
Comentarios
Great work! Now we need somebody to create the groups on iNaturalist ;-)!
Thanks!
Very good! I was looking for materials that lists the members of the species groups of Aradus.
Do you know any material for the species groups of east Palaearctic?
You would need to send an email to Ernst Heiss....he is our specialist on this group.
Thank you for the kind words!
Unfortunately, I only know of mentions of certain groups of the Eastern species. For example, the "transiens group" (omeiensis, quinlingshanensis, transiens, miyamotoi, malaisei, smetanai) distinguished by Ernst Heiss (https://www.zobodat.at/pdf/LBB_0035_1_0173-0180.pdf). But it seems to me that even the division of Western species needs to be clarified, taking into account species that so far remain outside any group. And before classifying the entire subgenus, I am inclined to consider any such division only preliminary. Therefore, in compiling the list, I limited the division to that of Heiss and Péricart.
P.S. A link to key to Korean species has been added.
Thanks, @kgrebennikov!
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